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Brightness spectrum 
of EGFP-mRFP1 (solid black) expressed in COS cells. Also shown is the expected apparent brightness spectrum 
for a sample composed of 65% EGFP and 35% of fluorescent EGFP-mRFP1 with a 0.52 FRET efficiency (gray solid). A single-species model (
) with 
(dashed black) fails to describe the experimental spectrum in its entirety.
