TABLE 2.
Changes in gene expression at different times during the Pi starvation response of C. glutamicum
ORF | NCBI no. | Annotation | Gene | Avg mRNA level after shift of Pi-limiting conditions/avg mRNA level before shift to Pi-limiting conditionsb
|
|||||
---|---|---|---|---|---|---|---|---|---|
10 min | 30 min | 60 min | 90 min | 120 min | 180 min | ||||
195 | Cgl2814 | Sulfate adenylate transferase subunit 1 | 2.2c | 0.3c | 0.2c | 0.4c | |||
312 | Cgl2917 | Putative integral membrane transport protein, similar to shikimate transport protein shiA | 10.0 | 0.4 | 0.6 | 0.5c | |||
348 | Cgl2946 | Hypothetical membrane protein | 1.2c | 0.7 | 0.3c | 0.2c | 0.2c | 0.1c | |
376 | Similar to heavy-metal-transporting ATPases | 2.2c | 5.7c | 3.4 | 2.5 | 1.3 | 1.0 | ||
378 | Cgl2965 | Two-component system, response regulator | 1.5 | 7.1c | 6.0c | 4.3c | 2.1c | 1.3 | |
380 | Cgl2966 | Hypothetical protein | 8.3c | 6.4c | 2.7c | 1.3 | 1.0 | ||
381 | Cgl2967 | Putative multicopper oxidase | cumA | 1.8c | 9.3c | 6.1c | 3.1c | 1.3 | 0.8 |
383 | Cgl2968 | Hypothetical protein | 1.8 | 5.7c | 3.4 | 1.0 | 0.6c | ||
399 | Cgl2979 | Hypothetical protein | 3.9c | 4.8c | 1.7 | 1.1 | 1.1 | 0.9 | |
403 | Cgl2982 | Single-stranded DNA-binding protein | ssb | 0.6c | 0.2c | 0.2c | |||
441 | Cgl0065 | Phosphate starvation-inducible protein | phoH | 1.1 | 1.8 | 3.3c | 4.4c | 4.7c | 4.5c |
442 | Cgl0065 | Phosphate starvation-inducible protein | phoH | 1.4 | 1.4 | 5.9c | 4.3c | 6.8c | 4.8c |
766 | Cgl0328 | 5′-Nucleotidase or related esterase | 1.2 | 7.3c | 6.5c | 19.3c | 10.3c | 7.2c | |
854 | Cgl0388 | Hypothetical membrane protein | 0.2c | 0.1c | 0.2c | 0.3c | 1.9 | 1.3 | |
856 | Cgl0390 | ABC-type transporter, permease component, similar to heme uptake systems | 0.4c | 0.2c | 0.3 | 0.5 | 1.6 | ||
857 | Cgl0391 | ABC-type transporter, ATPase component, similar to heme uptake systems | 0.3c | 0.2c | 0.3 | 0.5 | 1.5 | 0.9 | |
976 | Cgl0485 | Ribosomal protein L10 | 0.8 | 0.8 | 0.2c | 0.1c | 0.1c | 0.0c | |
978 | Cgl0486 | Ribosomal protein L7/L12 | 0.7 | 0.8 | 0.2c | 0.3c | 0.3 | 0.2c | |
988 | Cgl0493 | Ribosomal protein S12 | 1.5c | 0.8 | 0.3c | 0.3c | 0.2c | 0.2c | |
992 | Cgl0514 | Ribosomal protein S3 | 0.9 | 0.7 | 0.2c | 0.1c | 0.1c | 0.1c | |
993 | Cgl0515 | Ribosomal protein L16/L10E | 0.7 | 0.1c | 0.1c | ||||
994 | Cgl0516 | Ribosomal protein L29 | 0.9 | 0.7 | 0.2c | 0.1c | |||
999 | Cgl0521 | Ribosomal protein L14 | 0.3c | 0.2c | |||||
1000 | Cgl0522 | Ribosomal protein L24 | 1.2 | 0.8 | 0.2c | 0.2c | 0.1c | 0.1c | |
1020 | Cgl0539 | Ribosomal protein L18 | 1.0 | 0.7 | 0.2c | 0.2 | 0.2 | 0.1c | |
1044 | Cgl0560 | Translation initiation factor IF-1 | 1.1 | 0.4 | 0.3c | 0.4 | 0.2c | 0.4 | |
1047 | Cgl0562 | Ribosomal protein S11 | 1.1 | 0.7 | 0.2c | 0.2c | 0.2c | ||
1048 | Cgl0563 | Ribosomal protein S4 | 1.0 | 0.8 | 0.3c | 0.2c | 0.2c | ||
1050 | Cgl0566 | Ribosomal protein L17 | 1.0 | 0.7 | 0.3c | 0.2c | 0.2c | ||
1086 | Cgl0596 | Hypothetical protein | 10.8c | 18.8c | 17.6c | 15.0c | 8.9c | ||
1168 | Cgl0665 | Similar to vibriobactin utilization protein viuB | 0.1c | 0.3c | 1.3 | 0.8 | |||
1169 | Cgl0666 | ABC-type transporter, ATPase component, similar to iron uptake systems | 0.2c | 0.2c | 1.1 | 0.7 | |||
1170 | Cgl0667 | ABC-type transporter, permease component, similar to iron uptake systems | 0.1c | 0.2c | 1.0 | ||||
1173 | Cgl0669 | ABC-type transporter, periplasmic component, similar to iron uptake systems | 0.2c | 0.2c | 0.2c | 0.3c | 1.0 | 0.7 | |
1205 | Cgl0695 | Methylisocitrate lyase | prpB1 | 3.3c | 3.5c | 4.7c | 3.9c | ||
1206 | Cgl0696 | Methylcitrate synthase | prpC1 | 0.6 | 1.2 | 3.7c | 4.2c | 2.9c | |
1346 | Cgl0807 | Siderophore-interacting protein | 0.1c | 0.2c | 1.1 | ||||
1347 | Cgl0808 | ABC-type cobalamin/Fe3+-siderophore transport | 0.1c | 0.1c | 0.1c | 0.2c | 1.3 | 0.8 | |
1349 | Cgl0810 | ABC-type cobalamin/Fe3+-siderophore transport | 0.1c | 0.1c | 0.2c | 0.2c | 1.0 | 0.5 | |
1351 | Cgl0812 | ABC-type cobalamin/Fe3+-siderophore transport | 0.2c | 0.2c | 0.3c | ||||
1352 | Cgl0813 | ABC-type cobalamin/Fe3+-siderophore transport | 0.1c | ||||||
1420 | Similar to ribosomal protein L33 of E. coli (putative sequencing error) | 1.1 | 0.6 | 0.2c | 0.4 | 0.2c | |||
1421 | Cgl0869 | Ribosomal protein L28 | 1.0 | 0.8 | 0.2c | 0.2c | 0.2c | 0.2c | |
1558 | Cgl0982 | AraC-type DNA-binding domain-containing protein | 0.2c | 0.2c | 0.2c | 0.4c | 2.6 | 1.1 | |
1598 | Cgl1021 | Hypothetical membrane protein | 0.8 | 1.6c | 4.5c | 7.8c | 7.8c | 6.9c | |
1645 | Cgl1067 | Cysteine sulfinate desulfinase | 1.1c | 1.2 | 0.5c | 0.3c | 0.2c | 0.3c | |
1646 | Cgl1068 | Nicotinate-nucleotide pyrophosphorylase | 1.0 | 1.0 | 0.3c | 0.1c | 0.1c | 0.1c | |
1647 | Cgl1069 | Quinolinate synthase | 0.8 | 1.1 | 0.3c | 0.1c | 0.1c | 0.1c | |
1655 | Cgl1077 | 2-Polyprenyl-6-methoxyphenol hydroxylase | 4.6c | 1.6 | 0.9c | 1.0 | 1.0 | 0.9 | |
1722 | Cgl1139 | Methionine synthase II | 0.7 | 0.2c | 0.2c | 0.6c | |||
1855 | Cgl1248 | Siderophore-interacting protein | 0.4c | 0.3c | 4.3 | ||||
1875 | Cgl2996 | myo-Inositol-1-phosphate synthase | 0.8c | 1.3c | 0.8 | 0.4c | 0.2c | 0.5c | |
2146 | Cgl2035 | ABC-type transport systems, periplasmic protein, similar to iron(III) dicitrate uptake systems | 0.3c | 0.2c | 0.3c | 1.6 | 1.0 | ||
2168 | Questionable ORF | 1.6c | 1.1 | 0.3c | 0.2c | 0.2c | |||
2261 | Cgl2133 | Predicted Co/Zn/CD cation transporter | 0.8 | 1.2 | 2.0c | 2.7c | 3.5c | 4.8c | |
2262 | Cgl2134 | Dehydrogenase | 0.6c | 1.2 | 2.3c | 2.5c | 3.3c | 4.4c | |
2553 | Cgl1460 | ABC-type transporter, ATPase component | 1.0 | 2.8 | 3.0c | 5.0c | 2.7c | 2.3/PICK> | |
2554 | Cgl1459 | ABC-type transporter, permease component | 1.2 | 6.3 | 3.6c | 3.4c | 2.9c | 2.1 | |
2614 | Questionable ORF | 1.2c | 0.7 | 0.3c | 0.2c | 0.2c | 0.1c | ||
2624 | Cgl1399 | Arginine repressor | argR | 5.0c | 0.7 | 0.7c | 0.8 | 0.8 | 0.8 |
2625 | Cgl1398 | Ornithine carbamoyltransferase | argF | 5.5c | 0.6 | 0.6c | 0.6c | 0.7 | 0.7 |
2637 | Cgl1387 | Glycerophosphoryl diester phosphodiesterase | glpQ | 1.5 | 2.8c | 4.7c | 5.0c | 9.2c | 7.5c |
2638 | Cgl1386 | Glycerol-3-phosphate ABC-type transporter, ATPase component | ugpC | 1.1 | 4.2c | 3.9c | 4.3c | 4.5c | 4.5c |
2639 | Cgl1385 | Glycerol-3-phosphate ABC-type transporter, periplasmic component | ugpB | 2.2 | 12.9c | 22.8c | 18.7c | 17.2c | 14.5c |
2640 | Cgl1384 | Glycerol-3-phosphate ABC-type transporter, permease component | ugpE | 1.6c | 9.8c | 22.3c | 15.0c | 11.7c | 18.1c |
2641 | Cgl1383 | Glycerol-3-phosphate ABC-type transporter, permease component | ugpA | 2.1 | 15.1c | 16.8c | 15.0c | 11.0c | 12.5c |
2647 | Cgl1379 | Ribosomal protein L35 | 1.6c | 1.0 | 0.4c | 0.3c | 0.2c | 0.2c | |
2648 | Cgl1378 | Translation initiation factor IF3 | 2.0c | 1.1 | 0.3c | 0.2c | 0.1c | 0.2c | |
2779 | Cgl1281 | Co/Zn/Cd efflux system component | 3.8c | 4.7c | 1.9c | 1.0 | 0.6c | 0.7c | |
2831 | Cgl2607 | Two-component system, response regulator | 8.1c | 2.3 | 1.4 | 1.1 | 1.1 | 1.5c | |
2832 | Cgl2606 | Two-component system, sensory transduction | 4.7c | 1.7 | 1.0 | 0.8 | 0.8 | 0.9 | |
2849 | Cgl2594 | Predicted deacetylase | 4.7 | 2.0c | 2.1 | 2.1c | |||
2851 | Cgl2592 | Predicted extracellular nuclease | nucH | 4.8c | 19.5c | 31.7c | 25.8c | 16.9c | |
2871 | Cgl2575 | Phosphate ABC-type transporter, periplasmic component | pstS | 5.0c | 18.0c | 23.2c | 21.5c | 19.1c | 13.8c |
2873 | Cgl2574 | Phosphate ABC-type transporter, permease component | pstC | 4.4c | 19.3c | 23.6c | 22.8c | 20.9c | 19.1c |
2875 | Cgl2573 | Phosphate ABC-type transporter, permease component | pstA | 1.9 | 5.8c | 12.5c | 8.6 | 7.1c | 6.9c |
2876 | Cgl2572 | Phosphate ABC-type transporter, ATPase component | pstB | 2.7c | 9.0c | 12.4c | 14.5 | 10.3c | 8.3c |
2895 | Questionable ORF | 1.3 | 0.7 | 0.3c | 0.2c | 0.2c | 0.1c | ||
3081 | Cgl2336 | Hypothetical protein | 1.0 | 2.8c | 3.8c | 4.2c | 4.6c | 4.2c | |
3082 | Cgl2336 | Hypothetical protein | 4.3c | 5.3 | 5.0c | 5.3c | |||
3086 | Questionable ORF | 0.5c | 0.2 | 0.4c | 0.3c | 0.5c | |||
3326 | Cgl1067 | Cysteine sulfinate desulfinase | 0.6 | 0.2c | |||||
3347 | Cgl1714 | Hypothetical protein | 0.2c | 0.2 | 0.3c | 4.2 | 2.5 | ||
3362 | Questionable ORF | 1.8c | 2.7 | 4.7c | |||||
3442 | Cgl3064 | Hypothetical membrane protein, similar to phosphoesterases | 2.3 | 5.3 | 7.3 | 11.9c | 6.3 | ||
3458 | Cgl3075 | ABC-type transport systems, periplasmic component, similar to iron uptake systems | 0.1c | 0.1c | 0.1c | 0.1c | 0.5 | 0.3 | |
3472 | Cgl2526 | Ferritin-like protein | 10.3c | 10.3c | 5.8c | 5.5c | 1.8 | 1.6 | |
64185 | Cgl2389 | Acyl-CoA:acetate CoA transferase beta subunit | 7.3c | 1.5 | 0.4 | ||||
64187 | Cgl2393 | Predicted hydrolase/acyltransferase | 7.7c | 1.7 | 0.8 | ||||
66073 | Cgl2397 | Protocatechuate 3,4-dioxygenase beta subunit | 5.2c | 1.8 | 1.3 | 1.1 | 1.1 | 1.0 | |
66513 | Cgl2392 | Acetyl-CoA acetyltransferase | 4.4c | 1.8 | 0.6 | 0.5 | 0.7 | ||
69134 | Cgl2642 | Phosphotransferase system IIC component | 1.3 | 0.5 | 0.4c | 0.2c | 0.3c | 0.2 |
The relative mRNA levels 10, 30, 60, 90, 120, and 180 min after the shift to Pi-limiting conditions compared to the levels under preshift conditions are averages from three experiments. Only ORFs whose mRNA levels were equal to or greater than 4 or equal to or less than 0.25 at at least one time are shown.
Boldface type indicates relativemRNA levels that were equal to or greater than 10.
P < 0.05 as determined by a t test.