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Two kinds of events may occur backward in time in the genealogy of the linked neutral variant in the wild-type background: (I) coalescence events and (II) recombinations with a beneficial allele denoted by back-recombinations. (A) The average number of these events increases with the recombination rate ρ, but (B) for fixed ρ/α stays bounded for a wide range of values of the selection strength α. The Yule approximation does not allow for any of these events.
