LTP, basal synaptic transmission, and short-term plasticity in the
hippocampal CA1 region of WT and KO mice. (A) LTP was
induced by a repeated strong tetanization protocol consisting of 3
stimulus trains at 100 Hz (0.2 msec per polarity) with a 10-min
intertrain interval. During the initial phase of potentiation, WT mice
attained nearly the maximal level of potentiation after the first
tetanus, whereas KO mice progressively increased the potentiation after
each tetanus. For KO (but not WT) mice, the magnitude of the third
tetanus potentiation was significantly greater than that of the first
(P = 0.018 Wilcoxon matched-pairs signed rank
test). Arrows indicate the time of tetanization. (B)
Input/output curves of the KO and WT mice. fEPSP slopes were recorded
at increasing stimulation intensities until a maximum was attained.
There were no significant differences between the two groups.
(C) Short-term plasticity as evaluated by paired-pulse
stimulation at different IPIs. Although paired-pulse stimulation at an
IPI of 10 ms resulted in PPD, at all other IPIs, PPF appeared. Note the
stronger PPD of KO mice (P < 0.05).
(D) Decay of fEPSPs during the first tetanic
stimulation. The slope of 15 consecutive fEPSPs immediately at the
onset of tetanic stimulation was determined, averaged across each
group, and plotted vs. time. The curves were obtained by nonlinear fits
with a two-phase exponential equation. The fEPSPs of KO mice decayed
much faster than the WT in the initial part (first 100 ms) of the decay
(P = 0.022, ANOVA with repeated measures).
Insets show analogue traces of the first four fEPSPs
during the first tetanization. Note the marked decay of fEPSP slope of
the KO mouse between the first and second recording. (E)
Representative analogue recordings of fEPSP changes in a WT vs. KO
during the first tetanization. The faster decay of fEPSPs in the KO
results in a smaller depolarizing envelope evoked by summation of the
fEPSPs (area: WT, 878.4 units; KO, 530.5 units). The applied sampling
rate permits a clear distinction of the fEPSPs between the WT and KO.