Schematic neuroanatomy for area W and its input. The thick dashed line
separates area A from area W; the thin dotted line separates layers 2 +
3 from layer 4 in area W. Small filled circles indicate excitatory
connections, whereas small open circles indicate inhibitory
connections. The connections of a typical α-cell and a typical
β-cell, both shown in the center, are sketched. In the simulations,
area W is small, containing 325 neurons of each α- and β-type, and
a given cell makes synapses on 15–30% of these cells. Our simulation
contains 10 different γ-cells, each selective for a different
temporal pattern. Each γ-cell receives inputs from 30–80 cells of
each type, α and β.