t |
Time units (generally in years) |
N1
|
No. of age-1 individuals (newborns) produced in one time interval |
Nx |
No. of individuals of age x in the population |
∑Nx
|
Total no. of individuals in the population |
j |
Age at first maturity |
q |
Age at senescence |
N |
Population size, expressed as the total census size (), no. of reproductive adults (Nadult = ), or no. of newborns (N1) |
T |
Generation length (mean age of parents) |
bx |
Mean no. of offspring produced in one time interval by individuals of age x
|
lx |
Fraction of a cohort that survives to age x
|
sx |
Fraction of individuals of age x that survive to age x + 1 |
dx |
Fraction of individuals of age x that die before reaching age x + 1; dx = 1− sx
|
vx |
Reproductive value for individuals of age x
|
αx
|
Fraction of a cohort that reaches age x and then dies; αx = lxdx
|
λ |
Population growth rate per time period |
k |
No. of gametes contributed to the next generation by an individual over its lifetime |
|
Mean lifetime k for individuals that die between ages x and x + 1 |
σx2
|
Variance in lifetime k of individuals that die between ages x and x + 1 |
|
Mean lifetime k for all individuals breeding within a generation |
Vk |
Variance in lifetime k among all individuals within a generation |
Ne
|
Effective population size for a generation |
|
An estimate of effective size based on genetic data for two samples taken 1 or more time units apart |
|
Instantaneous effective size that describes the actual rate of change in the population at a point t in time |
S |
No. of individuals per time interval sampled for genetic analysis |
L |
Time interval between samples |
g |
Elapsed no. of generations between samples; g = L/T
|
F |
Standardized variance of allele frequencies |
Px(t) |
Allele frequency in age class x at time unit t
|
|
Population allele frequency at time t, weighting each individual equally |
|
Population allele frequency at time t, weighting each individual by its reproductive value |
C |
A constant that depends on life-history parameters of the population, used to estimate Ne in the temporal method of Jorde and Ryman (1995)
|