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(a) Schematic representation of the structure of aggrecan (contour length ∼400 nm), illustrating the core protein backbone (cp) and its three globular domains (G1, G2, G3), and the grafted chondroitin sulfate (CS) and keratan sulfate (KS) glycosaminoglycan (GAG) side chains. CS-GAG contour length ∼40 nm, intermolecular spacing ∼2–4 nm; N = N-terminal; C = C-terminal. (b) Disaccharide constituents of chondroitin-4-sulfate GAG and KS-GAG.
