Figure 1.

The model. (A) An input of intermediate contrast to the model. Each bar excites the principal model cells with the appropriate preferred locations and orientations. (B) The local principal (pyramidal) cells and interneurons are roughly reciprocally connected. Each pyramidal cell receives direct input from no more than one bar in the input in A. The pyramidal cells interact with each other (monosynaptically and disynaptically) via horizontal connections, and determine C—the output of the model, with the target’s z score indicated. The thicknesses of the bars are proportional to the temporal averages of pyramidal outputs.