The HD and CterC region confer AbdA activity in the epidermis. (A) Wild-type cuticle indicating the morphology of the thoracic (T1, T2, T3) and first two abdominal segments (A1, A2). Cuticular transformations associated with ubiquitous expression of Ubx (B), abdA (C), and chimera A (D). Ubx and abdA, respectively, transform segments anterior to their normal expression domains in A1 and A2 metameres. Chimera A phenocopies AbdA: it transforms the identity of thoracic and the first abdominal segments in an A2 identity.