Dicer is required for completion of meiotic maturation. (A) Schematic of oocyte maturation and early embryo development. The transition from fully grown GV oocyte to the late one-cell embryo occurs in the absence of significant transcription and relies on translation of maternal mRNA transcripts. The first part of this transition, when an oocyte completes meiosis I and arrests at metaphase of meiosis II is known as meiotic maturation. Zp3-Cre is expressed only in growing oocytes. (B) The maturation stage of Dicer−/− and control oocytes was scored by the presence of a GV (GV oocyte), a polar body (MII oocyte), or neither a GV nor a polar body (MI oocyte). Six-hundred-ninety-four Dicer+/+ and Dicer+/− and 582 Dicer−/− oocytes were analyzed in total. Error bars denote standard error of the mean. (C,D) Spindle morphology in Dicer−/− oocytes matured in vitro to metaphase I (C) or metaphase II (D). Oocytes were fixed in 3.7% paraformaldehyde and stained with a β-tubulin antibody; DNA was counterstained with DAPI or Sytox green. Representative images are shown. Bar, 25 μm. (E) Cortical actin and polar body morphology in mutant oocytes. Oocytes were fixed in 3.7% paraformaldehyde and stained with a β-tubulin antibody; F-actin was labeled with phalloidin-Alexa fluor 635 and DNA was counterstained with DAPI. The arrowheads indicate the position of the first polar body. β-Tubulin is often present inside the polar body in both wild-type and mutant oocytes.