Table 3.

Mean substitution parameters for Or/Gr genes and control genes along the D. sechellia and D. simulans lineages

Genes	Parameter	sechellia, mean	simulans, mean	Paired Wilcoxon P value
Ors	Ks	0.033	0.030	0.06
	Ka	0.005	0.003	<0.0001
	Ka/Ks	0.200	0.121	<0.0001
Grs	Ks	0.031	0.027	0.04
	Ka	0.007	0.004	<0.0001
	Ka/Ks	0.357	0.184	<0.0001
Ors+Grs	Ks	0.032	0.028	0.006
	Ka	0.006	0.003	<0.0001
	Ka/Ks	0.278	0.152	<0.0001
Controls	Ks	0.030	0.023	0.0001
	Ka	0.004	0.002	<0.0001
	Ka/Ks	0.145	0.117	0.001

The paired Wilcoxon P value tests the null hypothesis that the mean difference between the D. sechellia and D. simulans orthologs of each gene is zero.