R-cell differentiation and patterning appear normal in bks mutants. Wild-type (A, B, C, and D) and bks4 mutant (E, F, G, and H) third-instar eye-imaginal discs stained with cell-type-specific markers: anti-Prospero (A and E); anti-Boss (B and F); anti-Bar (C and G); and anti-Elav (D and H). Apical surface is up and posterior to the left. (Bar = 15 μm.)