Figure 4.

Schematic of interacting sub-populations and terms used to estimate the coupling. The rate of change of density in μ₂ over time, ${\dot{\mu }}_2$, depends on μ₁ owing to coupling. The density in population 1 leads to a firing rate, y₁, which modulates the synaptic channel-opening rate of population 2, parametrized by θ₂, which in turn modulates ${\dot{\mu }}_2$. The coefficient of coupling is calculated using the chain rule.