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Growth of wild-type P. aeruginosa PAO1 and swarming-deficient mutants in liquid BM2 swarm medium. Growth was measured at 37°C using a TECAN Spectrafluor Plus. Symbols: ▴, wild-type PAO1; ▪, PA5208 mutant; ×, folK mutant; ⧫, PA3817 mutant.