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. 2002 Nov;12(11):1679–1686. doi: 10.1101/gr.287302

Table 2.

Nucleotide Biasa at the Substitution Site and Immediate Adjacent Sites for Each Chromosome

Chromosome GC (%)b −1 0 +1





A (%) C (%) G (%) T (%) A (%) C (%) G (%) T (%) A (%) C (%) G (%) T (%)












 1 41.72 1.01 5.37 −1.59 −4.79 −4.61 4.50 4.53 −4.42 −5.03 −1.39 5.48 0.94
 2 40.14 2.07 4.55 −1.67 −4.95 −5.39 5.33 5.27 −5.22 −4.55 −1.61 4.65 1.51
 3 39.86 2.35 3.90 −1.74 −4.51 −5.50 5.43 5.34 −5.27 −4.26 −1.51 4.09 1.68
 4 38.26 2.49 3.91 −1.49 −4.90 −5.94 6.00 5.76 −5.82 −4.33 −1.35 3.86 1.81
 5 39.72 0.92 5.02 −1.39 −4.55 −5.47 5.25 5.41 −5.19 −4.58 −1.55 5.29 0.82
 6 39.62 1.19 4.88 −1.35 −4.73 −5.43 5.38 5.41 −5.36 −5.20 −0.98 5.22 0.97
 7 40.65 1.47 4.99 −1.78 −4.67 −5.38 5.30 4.95 −4.86 −4.55 −1.94 5.26 1.24
 8 40.07 2.58 3.40 −1.79 −4.20 −5.37 5.56 5.52 −5.71 −3.91 −1.85 3.40 2.35
 9 41.38 2.24 4.48 −2.27 −4.45 −5.11 5.22 4.73 −4.84 −4.16 −1.98 4.54 1.61
10 41.56 1.22 5.51 −1.74 −5.00 −4.75 4.72 4.42 −4.39 −4.86 −1.57 5.66 0.77
11 41.65 0.81 5.62 −2.55 −3.88 −4.45 4.54 4.50 −4.59 −3.68 −2.11 5.44 0.35
12 40.77 1.39 5.37 −1.84 −4.92 −4.94 4.91 4.98 −4.95 −4.68 −1.42 4.89 1.20
13 38.63 1.17 4.71 −1.32 −4.56 −5.64 5.63 5.57 −5.57 −4.78 −1.31 4.87 1.22
14 40.76 1.30 4.46 −1.88 −3.88 −4.77 4.74 4.84 −4.81 −3.38 −2.12 4.67 0.83
15 42.12 1.26 4.94 −1.88 −4.33 −4.82 4.66 4.62 −4.46 −4.25 −1.80 4.88 1.16
16 44.83 0.99 5.34 −1.84 −4.49 −4.09 3.79 3.87 −3.57 −4.61 −1.76 5.40 0.97
17 45.02 0.90 6.37 −2.43 −4.83 −3.61 3.62 3.00 −3.00 −4.59 −2.19 6.59 0.20
18 39.84 1.49 4.78 −2.16 −4.11 −5.20 5.12 5.39 −5.30 −4.06 −1.77 5.44 0.38
19 48.33 −0.42 7.51 −2.54 −4.57 −1.98 1.82 1.99 −1.84 −4.73 −2.63 7.79 −0.45
20 44.11 0.92 6.34 −2.02 −5.25 −3.31 3.54 3.62 −3.85 −4.91 −1.51 5.86 0.54
21 40.89 2.94 3.87 −2.27 −4.55 −5.13 5.04 4.52 −4.44 −4.48 −2.46 4.67 2.27
22 47.64 −0.40 7.74 −2.28 −5.06 −2.48 2.37 2.26 −2.15 −5.18 −2.09 7.87 −0.60
X 39.39 1.33 3.95 −0.88 −4.39 −5.41 5.24 5.83 −5.66 −4.85 −0.91 3.98 1.79
Y 39.11 3.18 3.02 −1.84 −4.37 −5.05 4.90 6.40 −6.25 −3.38 −0.87 3.85 0.40
Genome 40.90 1.43 4.91 −1.70 −4.62 −4.94 4.92 4.91 −4.88 −4.44 −1.59 5.05 0.99
a

Bias is expressed as a deviation from the chromosome-specific average. 

b

The average GC content on chromosome.