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| Symbol | Value for dispersal |
|---|---|
| f• | f |
| fp |  |
| f1 |  |
 |
(1 − d•)d•f + (1 − d•)2σ2 |
 |
sd•f |
 |
 |
 |
(1 − d1)d1f + (1 − d1)2σ2 + sd1f |
d•, dispersal rate of the offspring of the focal individual; 
average dispersal rate of the offspring of individuals from the focal deme (
); d1, average dispersal rate of the offspring of individuals from different demes
