TABLE 2.

Model parameter definitions

Parameter	Definition
	Viability of an AiAj female (f) or male (m) in which neither allele at the primary locus is imprinted or silenced.
	Viability of an AiAj female (f) or male (m) in which the maternal allele, Ai, is imprinted.
	Viability of an AiAj female (f) or male (m) in which the paternal allele, Aj, is imprinted.
	Viability of an AiAj female (f) or male (m) in which both alleles at the primary locus are imprinted.
μ	Rate at which mutation changes A1 to A2 when A1 is selectively advantageous in both females and males.
h, s	Dominance (h) and selection (s) coefficients in females.
k, t	Dominance (k) and selection (t) coefficients in males.
η, α, γ	Rate at which individuals with M1M1, M1M2, and M2M2 genotypes at the modifier locus imprint gametes in the trans-acting model.
ω, ν	Relative viability of females and males, respectively, with an imprinted allele when there is direct selection for imprinted expression in trans-acting model.