| Modifier linkage | Imprinting type | Resident modifier allele | Invading modifier allele | Jacobian matrix |
|---|---|---|---|---|
| Autosomal | Maternal | M1 | M2 | JAM1 |
| Autosomal | Maternal | M2 | M1 | JAM2 |
| X linked | Maternal | M1 | M2 | JSM1 |
| X linked | Maternal | M2 | M1 | JSM2 |
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