Table 4.
Details of study sites.
| Locality | Abbreviation | Coordinates | Origin |
| Madirovalo | madi | 16°22'45.6"S, 46°29'01.9"E | IRS 0 |
| Ampijoroa | jbb | 16°17'S, 46°48'E | IRS I |
| Mariarano | mari | 15°28'50.3"S, 46°41'19.0"E | IRS I |
| Tananvaovao | tan | 15°28'15.5"S, 46°39'59.4"E | IRS I |
| Ambodimahabibo | bibo | 15°29'54.2"S, 47°28'47.2"E | IRS II |
| Ambongabe | amb | 15°19'38.3"S, 47°40'44.4"E | IRS III |
| Anjiamangirana I | anji | 15°09'24.6"S, 47°44'06.2"E | IRS III |
| Ankarafa | sah | 14°22'47.8"S, 47°45'26.3"E | IRS IV |
| Mahilaka | mah | 14°17'12.0"S, 48°12'12.0"E | IRS V |
| Lokobe | lok | 13°23'23.9"S, 48°20'31.0"E | IRS VI |
| Manehoka | oka | 13°25'49.0"S, 48°47'51.0"E | IRS VI |
| Ankavana | anka | 12°46'55.7"S, 49°22'27.4"E | IRS VII |
| Ankarana | kar | 12°58'05.0"S, 49°08'18.0"E | IRS VII |
| Analabe | ana | 12°45'13.8"S, 49°30'03.9"E | IRS VII |
| Kirindy | kir | 20°03'S, 44°37'E | West |
| Mantadia | man | 18°47'S, 48°25'E | East |