Orthogonal niche axes. (A) Circular
niches of radius r randomly intersecting a habitat space
with orthogonal axes of lengths ar and
br. (B) Expected portion of habitat space
covered by species increases asymptotically with number of species in
the community. One-thousand trials were made for each value,
N, of species richness. In each trial, a set of
N circular niches of radius 0.218 were randomly placed
on a habitat space in which axes ranged from 0 to 1, and the portion of
the habitat space covered by this set of N species
determined. At least some part of the niche (circle, A)
of a species had to intersect the habitat space. •, means of
1000 trials. ·, 100 random trials for each value of N.
The solid curve is the theoretical value from Eq. 5.
(C) Variance in coverage declines to zero from a peak at
intermediate values of species richness.