Phylogenetic position of clones obtained in this study (shown in bold) affiliated with the Proteobacteria, inferred by maximum-likelihood analysis of 1,348 valid alignment positions. Greek letters represent the different subgroups. For details on tree calculation, evaluation, and notation, see the legend to Fig. 3. The tree was rooted with Thermus aquaticus (accession no. L09663), Aquifex pyrophilus (accession no. M83548), Thermotoga maritima (accession no. M21774), and Verrucomicrobium spinosum (accession no. X90515) as outgroup.