TABLE 2.

Relative mean bias and root relative mean squared error of the conventional G_ST and empirical Bayes estimators of pairwise F_ST from 1000 simulations under the stepping-stone models (see text)

	Relative mean bias		Root relative MSE
	Conventional	Empirical Bayes	Conventional	Empirical Bayes
0.001 (case 1)	1.165	1.239	2.352	1.817
0.0005 (case 2)	2.296	1.718	4.362	2.527

The number of subpopulations (K) was set at 15 and the pairwise F_ST between two adjacent populations was set at 0.001 (case 1) and 0.0005 (case 2). The sample size N_k/2 (individuals) was common to the localities and set at 20 individuals.