TABLE 3.

Average rate of fitness loss with a delta-function distribution of s-values

N	U	s	R1Da	Nebb	U2c	s2c	d
104	0.1	0.0006	0.060	6667	0.10	0.006	0.067
104	0.1	0.001	0.058	4000	0.099	0.001	0.062
104	0.1	0.003	0.039	1333	0.094	0.003	0.048
104	0.1	0.01	0.015	400	0.068	0.01	0.022
104	0.1	0.02	0.0031	200	0.022	0.02	0.0023

^aRatchet rate in the 1D Haigh model for the given values of N, U, and s.

^bEffective population size calculated as N_eb = 4/s.

^cEffective mutation rate and selection coefficient for the fraction of sites that are classified as category 2 as described in the text; the fraction of sites located in category 2 is U₂/U.

^dClick rate in the category 2 sites calculated from a 1D Haigh model with parameters N_eb, U₂, and s₂. When s approaches 0.0243, R^1D of the Haigh model is 0.01U and all sites would be located in category 1 giving U₂ =