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. 2007 Nov 21;104(49):19210–19213. doi: 10.1073/pnas.0707929104

Table 1.

Characteristics of the Ugalla USO digging sites

Site Site area,* m2 No. of holes Maximum diameter, mm Minimum diameter, mm Depth, mm Species Tool(s)
1 28 9 226 ± 68 (90–350) 207 ± 59 (90–300) 108 ± 47 (50–210) Dolichus kilimandscharicus
2 n/a 1 n/m n/m n/m Unidentified
3 7 4 n/m n/m n/m D. kilimandscharicus
4 n/a 1 280 220 180 Unidentified
5 6 4 116 ± 42 (60–155) 66 ± 20 (45–90) 55 ± 37 (30–110) Tacca leontopetaloides IS-S-001, IS-S-002, IS-W-001, IS-W-002
6 12 4 221 ± 107 (95–320) 171 ± 84 (65–260) 151 ± 62 (80–215) Raphionacme welwitschii IS-S-003
7 300 62 146 ± 41 (60–260) 115 ± 30 (60–200) 65 ± 29 (30–150) Brachystegia bussei, Smilax sp.
8 17.5 7 200 ± 82 (100–300) 154 ± 70 (50–250) 127 ± 76 (50–250) B. bussei
9 142.5 30 139 ± 52 (70–250) 110 ± 45 (50–230) 99 ± 53 (30–200) B. bussei, Smilax sp.
10 n/m >1 n/m n/m n/m B. bussei
11 277.5 96 97 ± 33 (40–180) 79 ± 27 (35–170) 81 ± 39 (30–190) Fadogia quarrei IS-B-001, IS-B-002

Data are ±SD with ranges in parentheses. n/a, not applicable; n/m, not measured.

*Minimum polygon area encompassing all holes identified at a site.

Although B. bussei does not have USOs sensu stricto, its young roots are used to store nutrients. Only the roots of very young saplings (up to 20 cm high) of this tree species were dug up; these taproots resembled in shape small carrots.

See Fig. 2 for an illustrated key to tool specimen catalog numbers.