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Combined effects of fertility selection and viability selection against a−, b−, and r− mutations on frequency of the r+ allele. C = 0.1, Mr = 10−8, X = 0.5, Mab = 0, quadratic selection against a− and b−, (see legend to Fig. 4). ○, Sr = 0; •, Sr = 0.1; □, Sr = 0.2; ▪, Sr = 0.4. Wr++ = Wr+− = 1, Wr− = 1 − Sr.
