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Analytically inferred and theoretically predicted numbers of charge-changing and charge-neutral substitutions.
| Charge-changing | Charge-neutral | Sum | |||
| Interior | Surface | Interior | Surface | ||
| Maximum likelihood-inferred numbers | 4 | 141 | 162 | 277 | 584 |
| Theoretical prediction | 35.92 | 133.45 | 184.46 | 230.17 | 584.00 |
| P value* | 0.00004 | 0.00150 | |||
* P values are from two-tailed exact tests.
