Path diagram of an AE Cholesky model for Fearless Dominance (FD) at Time 1 (T1) and Time 2 (T2). In this model, the variance at each time point is decomposed into additive genetic (A1, A2) and non-shared environmental effects (E1, E2). For ease of interpretation, shared environmental effects (c2) were omitted from this diagram. a11 and e11 = paths representing additive genetic and nonshared environmental contributions to the Time 1 phenotype, respectively; a21 and e21 = paths representing additive genetic and nonshared environmental contributions from Time 1 to the Time 2 phenotype, respectively; a22 and e22= paths representing additive genetic and nonshared environmental contributions unique to the Time 2 phenotype, respectively. These paths are squared to estimate the proportion of variance accounted for by additive genetic and nonshared environmental influences.