Data are consistent with I-Plat cell 71 exciting I-Aug cell 43, and E-Dec cell 46 exciting E-Aug cell 44 and inhibiting phrenic motoneurones. A, events indicated by the circled numbers in the designated cough period are detailed in the text. B, CCH; offset peak with a lag of 1.5 ms; HW, 3.0 ms; DI, 5.3; S, 0.41; 19082 reference and 4702 target spikes. C, CCH; offset peak with a lag of 1.5 ms; HW, 3.0 ms; DI, 3.2; S, 0.43; 27822 reference and 1938 target spikes. D, STA; number of trigger events, 32500. E, summary circuit of inferred connectivity.