Effects of breeding stage on singing variables and approach; bars show means ±SE. Breeding stages and sample sizes are: none (no nest) n = 7; build (nest building) n = 7, inc (incubating eggs) n = 9; nestl (feeding nestlings) n = 6; fledg (feeding fledglings) n = 5. (A) Matching rate as a function of breeding stage, contrast between build-incubate-fledgling stages versus none-nestling stages significant at P = 0.004 (Table 2). (B) Overlapping rate as a function of breeding stage. (C) Versatility as a function of breeding stage, contrast between fledgling versus all other stages significant at P = 0.0004 (Table 2). (D) Approach score as a function of breeding stage, contrast between fledging versus all other stages significant at F1,29 = 5.41, P = 0.027.