‘Inter’ indicates interneurones, whereas ‘output’ stands for output neurones. A, MNSI and LICI are independently mediated and do not interact; B1, MNSI and LICI are mediated simultaneously via the same interneurones; B2, MNSI is sequentially mediated via the LICI; C, MNSI and LICI are independently mediated, but interact via excitatory or inhibitory mechanisms.