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PMFs for the ionizable residues: (A) Arg, (B) Lys, (C) Glu, (D) Asp. The PMFs for the ionized forms have been set to zero in the water phase. The neutral PMFs have been offset by the free energy to neutralize the residue in bulk water at pH 7.0. Error bars indicate the SE based on the asymmetry between the two leaflets of the bilayer.
