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. 2006 Oct 3;4(12):143–153. doi: 10.1098/rsif.2006.0167

Table 2.

Symbols used.

S proportion susceptible to both strains
Ii proportion in primary infection with strain i={1,2}
Ri proportion immune to strain i only
Ji proportion in secondary infection with strain i={1,2}
R proportion immune to both strains
V proportion vaccinated and still immune to strain 1
IV2 proportion vaccinated and infected by strain 2
β rate of transmission
γ rate of recovery
μ birth and death rate
p vaccine coverage (proportion of newborns vaccinated)
σ rate of loss of natural immunity
σV rate of loss of vaccine immunity
θ cross-protection conferred by primary infection (reduction in susceptibility)
ν cross-protection conferred by primary infection (reduction in infectious period)
τ cross-protection conferred by vaccination (reduction in susceptibility)
η cross-protection conferred by vaccination (reduction in infectious period)
R0 basic reproductive ratio of the pathogen: R0=β/(γ+μ)