Nodule development (Nod) in white clover 40 days after inoculation with the wild-type ANU843 (A, C, and E) or the CelC2− mutant ΔcelC2 (B, D, and F); shown are portions of nodulated roots from inoculated plants (A and B), longitudinal sections of the induced nodules (C and D), and whole-plant phenotypes (E–G). (Scale bars: 200 μm.)