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. 2007 Dec 19;104(52):20672–20677. doi: 10.1073/pnas.0705527105

Table 4.

Aggregate land use pattern measures

Pattern measure Developed land use pattern
Undeveloped land use pattern
All development
High-density residential
Low-density residential
1973 2000 % change 1973 2000 % change 1973 2000 % change 1973 2000 % change
Patch density 0.14 0.33 132.0 0.04 0.11 157.2 0.12 0.33 172.9 0.03 0.06 86.5
Mean patch size 0.60 0.52 −14.3 0.94 0.56 −40.4 0.27 0.25 −4.3 25.74 12.24 −52.4
Mean perimeter-to-area ratio 13.23 15.02 13.5 11.31 14.46 27.8 13.68 16.10 17.7 13.78 17.27 25.4
Contrasting edge ratio* 0.51 0.81 60.5 0.13 0.16 25.8 0.53 0.86 63.1 0.03 0.09 177.8
Contrasting edge proportion* 0.22 0.35 54.9 0.06 0.08 37.2 0.21 0.35 63.6 0.03 0.06 138.8
Mean dispersion* (1 km2) 0.37 0.38 2.7 0.23 0.21 −10.1 0.57 0.53 −6.3 0.03 0.08 131.9
Mean dispersion* (25 km2) 0.52 0.50 −3.0 0.39 0.33 −15.4 0.70 0.65 −7.2 0.07 0.12 62.5

*For developed (residential) land use measures, focal land use = developed (residential), and contrasting land use = undeveloped; for the undeveloped land use measures, focal land use = undeveloped, and contrasting land use = developed. Refer to Table 3 for pattern measure definitions.