Summary of collicular circuitry in the tree shrew (24). Cells in the lower SGS (a) project to the optic layer where they contact either optic layer cells (b) or apical dendrites of premotor cells in upper SGI (c). Optic layer cells project to the optic layer (SO) as well as to the adjacent layers. Premotor cells in lower SGI (d) receive input from either SO or upper SGI cells. l, lower; SGI, stratum griseum intermedium or intermediate gray layer; SGS, stratum griseum superficiale or superficial gray layer; SO, stratum opticum or optic layer; u, upper.