vertebrates, invertebrates (oligochaetaes, polychaetaes, crustaceans, molluscs, ants, termites), plant roots |
bioturbators, ecosystem engineers |
soil alteration, soil structure, organic matter and microbes mixing laterally and vertical depths |
plant roots, algae |
primary producers |
create biomass, stabilize soils and sediments, provide C and N at depth |
millipedes, centipedes |
shredders |
fragment, rip and tear organic matter |
bacteria and fungi, Collembola, mites, nematodes, rotifers, tardigrades, protozoa |
primary decomposers and secondary decomposers |
recycle nutrients, increase nutrient availability for primary production |
symbiotic (e.g. Rhizobium) and asymbiotic bacteria (e.g. Cyanobacter, Azobacter) |
nitrogen fixers |
biologically fix atmospheric N2
|
methanogenic bacteria, denitrifying bacteria |
trace gas producers |
C, N2, N2O, CH4 transfer and denitrification |
roots, soil organisms |
CO2 producers |
respiration, emission of CO2
|