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Typical positions of ions in the filter relative to pore-lining carbonyl oxygens (red spheres) from the indicated residues. Potassium ion is shown as a purple sphere, sodium ion as a green sphere. Illustrated in panels A–C are, respectively, a K+ ion in site S3 during inwards conduction, a Na+ ion and two water molecules during inward conduction, and a K+ ion and water molecules at the top of the filter pocket during outward conduction.
