| m | the anopheline density in relation to man, |
| a | the average number of men bitten by one mosquito in one day, |
| b | the proportion of those anophelines with sporozoites in their glands which are actually infective, |
| p | the probability of a mosquito surviving through one whole day, and |
| r | the proportion of affected people, who have received one infective inoculum only, who revert to the unaffected state in one day. |