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Structure of a chondroitin sulfate A-binding DBL3X domain. A, the var2csa protein consists of six extracellular DBL domains followed by a single transmembrane helix and a cytoplasmic acidic terminal sequence (ATS). Residues 1218–1577 form the DBL3X domain. B, orthogonal views of the DBL3X domain with subdomain 1 in yellow, subdomain 2 in orange, and subdomain 3 in red. C–E, subdomains 1, 2, and 3, respectively, with residues involved in disulfide bonds highlighted in green.
