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Superposition of three structures of HD-domain proteins with divalent cations and substrates (or natural inhibitors) bound. YfbR E72A complexed with Co2+ and TMP (this work) is shown in blue. S. equisimilis RelA complexed with Mn2+ and guanosine-5′-diphosphate-2′,3′-cyclic monophosphate is shown in magenta7. T. thermophilus dNTPase complexed with Mg2+ is shown in green8.
