Figure 4.—

Partial complementation among specific Doa alleles leads to strong female-to-male somatic sex transformations. (A and B) Wild-type female genitalia, light and scanning electron microscopic views. (C and D) Wild-type male genitalia. (E–G) An XX, Doa^E786/Doa^EMS2 female: (E) lateral view, note the male “clasper” (arrow) but typical female pigmentation; (F and G) ventral view, symmetrical male claspers (arrows), as well as the female genital sex comb (arrowheads), produce a doublesex-like phenotype. (H–K) Doa^EMS3/Doa^HD female: (H) lateral view, note the male pigmentation; (I and J) ventral view, genitals are rotated but nontransformed; (K) perfectly formed male sex comb on a prothoracic leg (arrow).