TABLE 3.

Summary of phenotypes observed in Doa heteroallelic combinations

1. Eyes	a. Rough, slight ommatidial disorganization
	b. Mosaic, nonuniform color
	c. Grossly disorganized ommatidia
2. Bristles: scutelar, postalar, dorsocantral, head, or other	a. Duplicated
	b. Missing
	c. Forked
	d. Misoriented
3. Wings	a. Additional or incomplete cross-veins
	b. Additional or incomplete longitudinal veins
	c. “Fragile,” damaged
	d. Heavy, blackened veins
	e. Held out
	f. Weak curly-like phenotype
4. Sex transformation	a. Female vaginal bristles nonsymmetric, missing
	b. Male clasper(s) on female vaginal plate
	c. Sex comb on female prothoracic legs
	d. Male genitals rotated
5. Aberrant sex ratio in F1	a. ≥3× female/male F1
	b. ≥3× male/female F1
6. Deformed legs	a. Metathoracic
	b. Mesothoracic
7. Substantial death during pupation
8. Nonreciprocal results (Table 4)