Each swrA promoter is sufficient for swarming. (A) Swarming ability in swrAA+ (PB5392) and swrAA mutant (PB5396) strains. (B) Swarming ability in swrAA+ strains carrying mutations in the putative σA-dependent swrA promoter (A−, PB5393), in the σD-dependent swrA promoter (D−, PB5394), and in both promoters (DA−, PB5395).