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The proposed mechanism for Cu(I) transfer between HAH1 and MNK4. A possible four-coordinate intermediate that would reside between the two three-coordinate states in the middle has been left out. In the HAH1 homodimer used as a mimic of the HAH1:MNK4 heterodimer in MD simulations, C14 and C17 of the target domain are replaced by Cys 12B and Cys 15B, respectively.
