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Linking number of episomal DNA molecules is altered by an act3 mutation. DNA was isolated from logarithmically growing yeast cells containing the YCp50 plasmid, electrophoresed in two-dimensional chloroquine-agarose gels, and subjected to Southern blot analysis with 32P-labeled URA3 DNA fragment. Left panel, DY3462 (wild-type); right panel, DY3189 (act3-3). The directions of the first and second dimensions are indicated.
