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Ribbon representation of the head region of chicken skeletal myosin in the nucleotide-free state (PDB 2mys; (1)) bound to an actin filament in the absence of ATP (2), showing the lever and hook axes, and the tilt (β) and twist (γ) angles. Pairs of cysteine residues introduced into the ELC and cross-linked using BSR-I2 (inset) are shown as spheres joined by rods.
