Top: Schematic illustration of long-distance links connecting the cortical areas related to language and action. Motor regions in red, inferior frontal (including Broca's) language area in purple, superior temporal (including Wernicke's) language area in blue. Bottom: A neurobiological model of language–action links: Cell assemblies processing specific language elements, for example action words, may bind information about word forms, which is laid down in perisylvian language cortex, with information about actions, which is laid down in different parts of the motor system (after Pulvermüller, 2005). Abbreviations: A1 = auditory “core” cortex, BPO = Broca, pars opercularis, BPT = Broca, pars triangularis, M1 = motor cortex, PMC = premotor cortex, PFC = posterior prefrontal cortex, WB = Wernicke, auditory belt area, WPB = Wernicke, auditory parabelt area.