(a,b) Early ommochrome development (cyrbia, himera, respectively) and (c,d) late melanin development (cyrbia, himera, respectively). Quantitative RT-PCR comparison of (i–iv) vermilion and (v–viii) cinnabar transcript abundance during pigment development in (i,v,iii,vii) H. e. cyrbia and (ii,vi,iv,viii) H. himera. Transcription levels are presented relative to control gene expression and error bars represent the standard deviation. Synthesis of red pigment occurs when the transcription of both genes spatio-temporally coincides in the forewing band (i,v).