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Top five 4-mer motifs in different species arranged in order, highest on the top
| Rank | Drosophila | Human | Mouse | Plant | Rat |
| 1 | TTTT | GCGG | AAAA | AAAA | GGAG |
| 2 | AAAA | GGCG | GGGG | TTTT | GGGG |
| 3 | ATTT | GGGG | CAGG | AAAT | AGAG |
| 4 | AAAT | GGGC | CCAG | TATA | CAGG |
| 5 | AATT | GCCC | GGGC | ATAT | CAGC |
