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Estimated reproductive fitness of five male colour morphs in two mating habitats. Morphs vary in mean reproductive fitness (error bars are ±1 s.e.) between and within beach (filled circles) and root (open triangles) mating habitats. Within a habitat different letters indicate morphs that are significantly different from each other, as determined by Tukey's HSD (qbeach=2.94, qroot=2.94, α=0.05).
